ICHNOTAXONOMY
OF THE FOSSIL FOOTPRINTS OF THE CONNECTICUT VALLEY (EARLY JURASSIC,
NEWARK SUPERGROUP, CONNECTICUT AND MASSACHUSETTS)
Emma Clare Rainforth
Submitted in partial fulfillment of the requirements for the degree of
Doctor of Philosophy in the Graduate School of Arts and Sciences
COLUMBIA UNIVERSITY
© 2005
Emma Clare Rainforth
All Rights Reserved
Abstract
From 1836 to 1865, Edward Hitchcock
named 94 ichnogenera and 216 ichnospecies (including unjustified
emendations), in a total of 269 combinations. After a comprehensive
examination of all syntypes, ichnospecies, and ichnogenera, tracing
both objective and subjective synonymies, 71 ichnogenera and 185
ichnospecies are valid. It is likely that this number can be reduced
significantly in the future by subjective synonymization. Type
ichnospecies are designated for six ichnogenera not previously
typified. Lectotypes are designated for 75 ichnospecies, all of which
had previously (C. H. Hitchcock 1865; Cushman 1904; Lull 1904a, 1915,
1953) had lectotypes invalidly designated. Five neotypes are designated.
Hitchcock’s (1836-1865) classification of Ornithichnites into Leptodactyli
and Pachydactyli (slender- and thick-toed respectively) is considered
to be a taphonomic rather than biologic division. For trackmaker
identification and subsequent biostratigraphic analysis, the
Pachydactyli are more useful. The ornithischian Pachydactyli have
recently been re-evaluated (Olsen and Rainforth 2003); the theropodan
Pachydactyli (Eubrontes ispp.)
require major revision and are expected to be valuable for
high-resolution biostratigraphy in the Newark Supergroup.
Otozoum moodii (Hitchcock
1847) is one of the classic Connecticut Valley ichnotaxa. It is
redescribed, and AC 4/1a re-established as the holotype. O. minus and O. caudatum are synonymized with O. moodii. The syntypes of Kalosauropus pollex are described
as O. pollex sp. nov.; K. masitisii is considered a nomen dubium. Cladistic,
quantitative, and comparative methods of trackmaker identification
suggest that Otozoum was a
prosauropod print. In quantitative analyses, only those phalanges
likely to contribute to footprint morphology were considered; claws
were excluded owing to their variable impression. Several diagnostic
characters distinguish Otozoum
from Brachychirotherium, Chirotherium, Batrachopus, Tetrasauropus, and Pseudotetrasauropus; these
ichnotaxa were probably made by crurotarsans. Otozoum is found primarily in
Hettangian strata of the Hartford, Deerfield, and Fundy basins (eastern
North America); it is also present in the Early Jurassic Navajo
Sandstone (Colorado Plateau, USA) and Clarens Formation (Lesotho). It
is unknown in older strata; Triassic material previously referred to Otozoum (including O. grandcombensis) instead belongs
in Pseudotetrasauropus.
Correlation of sediments and comparison of faunal assemblages,
particularly within the Newark Supergroup, suggest that the Otozoum trackmakers were restricted
to quite arid environments.
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Front
matter
Chapter 1: Introduction to Connecticut Valley
Ichnotaxonomy
Chapter 2: Status of the Ichnogenera
Ornithichnites,
Ornithoidichnites and
Eubrontes
Chapter 3: Connecticut Valley Vertebrate
Ichnogenera Containing Ichnospecies Named Prior to 1845 and not
Referred to
Eubrontes
Chapter 4: Connecticut Valley Ichnogenera Based on Ichnospecies Named
After 1845
Chapter 5: Status of Connecticut Valley Ichnotaxa Attributed to
Non-vertebrates
Chapter 6: Ichnotaxonomic Summary and Future Studies
Chapter 7: The Early Jurassic Ichnogenus
Otozoum
(see also Rainforth
2003)
Appendices (1 - Unpublished Amherst College
catalogs; 2 - Identity of figured specimens (1836-1865); 3 -
Otozoum; 4 -
Thenaropus)
References