ESTIMATING SPEEDS OF DINOSAURS FROM TRACKWAYS: A RE-EVALUATION OF
ASSUMPTIONS
Rainforth, Emma C., and Manzella, Melissa, Environmental Science,
Ramapo College of New Jersey, 505 Ramapo Valley Road, Mahwah, NJ 07430;
erainfor@ramapo.edu, mmanzell@ramapo.edu
For 30 years, using equations determined by Alexander (1976),
paleontologists have estimated speeds of dinosaurs from their
trackways; the fundamental assumption, based on limited observations,
was that dinosaur hip height is approximately four times foot length.
Thulborn (1990) subsequently determined that the leg length to foot
length ratio ranges from 4.5-6.0, according to type and size of
dinosaur. Given that the focus of many published footprint studies is
to estimate dinosaur speed from Alexander’s equations, it is critical
that the hip height of the track maker can be ascertained from the
footprint measurement. If there is high variability in the ratio of hip
height:foot length within a group of dinosaurs, speed estimates are
going to be extremely unreliable.
This study examined the relationship between foot length and leg length
in a variety of dinosaurs. Measurements of 24 dinosaur specimens
included ornithischians (bipedal and quadrupedal), theropods,
dinosauriforms, birds, and sauropods. Leg length was defined as femur +
tibia + metatarsal III; foot length was defined as that part of the
foot preserved in footprints (digit III phalanges ± claw +
metatarsal III – metatarsal IV). Leg and foot lengths were compared and
it was found that in bipedal ornithischians the foot to leg length
ratio is ~5.4-5.9, and in theropods, 2.8-4.2. Contrary to previous
studies, we found that (1) there is no correlation between dinosaur
size and foot to leg length ratio, (2) the foot to leg length ratio is
greater than previously estimated, and (3) the foot to leg length ratio
is highly variable for each group of dinosaurs. We conclude that
estimating dinosaur speed from trackways should not be undertaken
lightly, and the results be interpreted with caution.
Geological Association of New Jersey 24th Annual Meeting, October
12-13, 2007. In Rainforth, E.C. (ed.), 2007, Contributions to the
Paleontology of New Jersey (II): Field Guide and Proceedings,
Geological Association of New Jersey 24th Annual Conference and Field
Trip, p. 4.